Indeed, Tony. And I think, Chris, that if you look more widely at the science you’ll find that (as C&B note) the Phylogenetic Species Concept (PSC) WOULD take in such factors as ‘topography, *food, hunting, behaviour, breeding, and vocalisations*’ (as Tim says) in defining species. As this would cause many more species than we currently have (the number depending upon the defining characteristics applied by the PSC splitter), methods of species definition remain open to discussion. Thus C&B appear to follow a course of amending the BSC (Biological Species Concept, which incorporates their palaeological skills) with molecular/DNA research discoveries.
C&B would, I’m sure, be the last to want to stifle discussion on this interesting and constantly evolving topic.
Judith. L-A